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Looks like the mic for the Bluetooth phone prep, at least that's what it is on my Roadster. There's lots of similarities between the two cars. I would expect that they share the upper switch panel and dome light due to the raked windshields. The mic is up there on the hardtops and Clubmen, but I think the Roadster and Coupe get the mic on the steering column like the Convertible.
Not at all certain but it looks like a speaker\microphone used by Bluetooth\hands free. Is that a possibility. My '08 has the speaker in the overhead switch area and it isn't real effective.
That's where the knock-out gas is sprayed from when you press the wrong button in a Mini equipped with the 'James Bond' baddie' option (JB007). Press the grille three times in quick succession to operate the passenger ejection seat.